Hermann Bragulla

Hermann Bragulla

Associate Professor

Department of Comparative Biomedical Sciences

LSU School of Veterinary Medicine
Louisiana State University
Baton Rouge, LA 70803



Dr.habil, University of Berlin

PhD, University of Munich

DVM, University of Munich

Research Interest

Pre- and post-natal developmental processes of skin and skin modifications in terrestrial and marine mammals such as hooves and baleen, comparative morphology in reptiles, birds, and mammals 

Teaching Interest

Comparative morphology, i.e. anatomy, histology, and embryology of vertebrates, especially mammals and birds 

Clinical Interest

Age-related versus pathological morphological changes in mammals such as the temporomandibular joint in horses

Awards & Honors

2013, Teaching Award


Cruz-Sanabria, J. A., l. Gaeschen, H. H. Bragulla, M. Mitchell, and B. S. Leise (2021): A study of ultrasound-guided perineural injection of the caudal cervical spinal nerve roots in equine cadavers. Vet. Anaesth. Analg., 48 (4), 603-611

Schoenfuss, H. L., H. H. Bragulla, J. Schumacher, W. G. Henk, J. C. George, and D. J. Hillmann (2014): The anatomy of the larynx of the Bowhead whale, Balaena mysticetus, and its sound-producing functions. Anat. Rec., 297, 1316-1330

Levy, M., L. Gaschen, N. Rademacher, and H. Bragulla (2014): Technique for ultrasound-guided intraarticular cervical articular process injection in the dog. Vet. Radiol. Ultrasound, 55, 1-6

Middleton, G., D. J. Hillmann, J. Trichel, H. H. Bragulla, and L. Gaschen (2012): Magnetic resonance imaging of the ligamentous structures of the occipitoatlantoaxial region in the dog. Vet. Radiol. Ultrasound, 53(5), 545-551

Gaschen, L., A. Le Roux, L., Riggs, J. Trichel, H. H. Bragulla, N. Rademacher, and D. Rodriguez (2011): Magnetic resonance imaging in foals with septic arthritis.  Vet. Radiol. Ultrasound, 52(6), 674-705

Bragulla, H. H., and D. G. Homberger (2009): Structure and functions of keratin proteins in simple, stratified, keratinized and cornified epithelia.  J. Anat., 214, 516-559

Homberger, D. G., K. Ham, T. Ogunbakin, J. A. Bonin, B. A. Hopkins, M. L. Osborn, I. Hossain, H. A. Barnett, K. L. Matthews III, L. G. Butler, and H. H. Bragulla (2009): The structure of the cornified claw sheath in the domesticated cat (Felis catus): Implications for the claw shedding mechanism and the evolution of cornified digital end organs.  J. Anat., 214, 620- 643

Bragulla, H. H., and D. G. Homberger (2007): The role of the specific, profilaggrin-containing keratohyalin granules in the developing epidermis of the fetal horse hoof.  Pferdeheilkunde, 23(1), 5-20

Hirschberg, R. M., and H. H. Bragulla (2007): Funktionelle Aspekte der Angioarchitektur des Pferdehufs. [Functional aspects of the angioarchitecture in the equine hoof.]  Pferdeheilkunde, 23(1), 27-38

Schell, H., J. Lienau, D. R. Epari, P. Seebeck, C. Exner, S. Muchow, H. Bragulla, N. P. Haas, and G. N. Duda (2006): Osteoclastic activity begins early and increases over the course of bone healing.  Bone, 38(4), 547-554 


July 2011 – June 2012 EHSP-Grant; PI: Dr. Hermann H. Bragulla, co-PI: Dr. Mandi Lopez; Topic Keratin proteins change in chronic laminitic equine hooves

July 2008 – June 2010 EHSP-Grant; PI: Dr. Hermann H. Bragulla, co-PI: Dr. Ashley Stokes; Topic: Keratin proteins in the horse hoof.